The study reported that the main reason for the "alarming" decline in the world's mammals, birds, amphibians, reptiles and fish was the destruction of their natural habitats.

"Global patterns of rising extinction risk are most marked in Southeast Asia, where agricultural expansion, logging and hunting are the primary forces behind accelerating extinction rates," a summary of the study said.

The study, by 174 scientists around the world, was described as the first time the rate of decline among vertebrate species had been quantified on a global scale.

It was based on research into 25,000 species on the International Union for the Conservation of Nature's (IUCN's) "Red List" of threatened species.

IUCN members were due to discuss the findings of the research on Wednesday on the sidelines of a United Nations summit being held in the Japanese city of Nagoya to try to map out a plan to slow or stop the decrease in the world's biodiversity.

The summit is being held amid the backdrop of warnings from scientists that humans' destruction of nature is causing plant and animal species to become extinct at up to 1,000 times the natural rate.

The IUCN said last year the world was experiencing its sixth mass extinction in history, the last one being 65 million years ago when dinosaurs were wiped off the planet.

However the study published Wednesday said species loss and decline would have been 20 percent worse in the absence of conservation efforts to protect threatened species.

"Thus, while current conservation efforts remain insufficient to offset the main drivers of biodiversity loss... targeted conservation efforts have had a measurable positive impact on the planet's vertebrate species," the study said.

Some of the conservation strategies the study highlighted as being beneficial were captive breeding programmes, legislation to limit hunting, establishing protected areas and efforts to remove invasive alien species.

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